SciELO - Scientific Electronic Library Online

vol.16 issue2Evolución geomorfológica de Punta Rasa, Pcia. de Buenos Aires, Argentina author indexsubject indexarticles search
Home Pagealphabetic serial listing  

Services on Demand




  • Have no cited articlesCited by SciELO

Related links


Revista del Museo Argentino de Ciencias Naturales

On-line version ISSN 1853-0400

Rev. Mus. Argent. Cienc. Nat. vol.16 no.2 Ciudad Autónoma de Buenos Aires Dec. 2014



Cactaceae type specimens at BA. Argentinian Natural Sciences Museum Herbarium, Buenos Aires, Argentina


Valentin D. Fadel, Eugenia C. Alvarenga & Mirta O. Arriaga

BA Herbarium, Vascular Plants' National Colletion, Argentinian Natural Sciences Museum "Bernardino Rivadavia", Av. Ángel Gallardo 470, C1405DJR, Ciudad Autónoma de Buenos Aires, Argentina. e-mail:


The Herbarium from Argentinian Natural Sciences Museum "Bernardino Rivadavia" (BA Herbarium) was the first official herbarium in this country. It was initiated in 1853 and holds about 150.000 herbarium specimens at present, most of them collected all over Argentina, but also obtained by exchange with other national and international botanical institutions. About 100.000 correspond to vascular plants, and 800 represent nomenclatural type specimens. Cactaceae type material deposited at BA is presented. This article includes a list of the 60 type specimens alphabetically arranged; scientific names with taxon authors, original publication data and type category are transcribed from the original labels. Current accepted scientific names are also included. Under observations, ecological information or additional comments written by the author of the description are given. Under notes some consideration of our own, most regarding to the current synonymy, are given. These type specimens are kept in special metal cabinets, separated from the rest of the herbarium material and documented with a copy of the original diagnosis or publication. As part of the Digitalization of Biological Collections Project carried out by the museum, the complete information of each specimen was digitalized and is available online through the web site of the Sistema Nacional de Datos Biológicos (SNDB) and the Global Biodiversity Information Facility (GBIF).

Key words: Type specimens; Cactaceae; Herbarium BA; Argentina.


Ejemplares tipo de Cactaceae depositado en BA. Herbario del Museo Argentino de Ciencias Naturales "Bernardino Rivadavia", Buenos Aires, Argentina. El Herbario del Museo Argentino de Ciencias Naturales "Bernardino Rivadavia" (Herbario BA) fue el primer herbario oficial de este país. Iniciado en el año 1853, alberga actualmente unos 150.000 especímenes de herbario, la mayoría de ellos colectados a lo largo y ancho de Argentina pero también obtenidos por intercambio con otras instituciones botánicas tanto nacionales como internacionales. De todos ellos, aproximadamente 100.000 corresponden a especímenes de plantas vasculares y 800 representan especímenes tipo. Continuando con la publicación de los datos de los ejemplares tipo depositados en BA, se presenta un catálogo del material tipo de Cactaceae. El presente artículo incluye un listado de los 60 especímenes tipo alfabéticamente ordenados; se transcribe para cada uno de ellos el nombre científico y autor del taxón, así como la información original de etiqueta. En las observaciones se incluye la información ecológica o comentarios adicionales redactadas por el autor de la descripción. En cuanto a las notas provistas por los autores de este trabajo, se incluyen algunas consideraciones acerca de la sinonimia actual. Los ejemplares tipo son guardados en gabinetes metálicos especiales, separados de resto de material del herbario y documentados con una copia de la diagnosis original o su publicación. Como parte del Proyecto de Digitalización de las Colecciones Biológicas llevado a cabo por el museo, la información completa de cada especimen fue digitalizada y se encuentra disponible a través del portal del Sistema Nacional de Datos Biológicos (SNDB) y la Global Biodiversity Information Facility (GBIF).

Palabras clave: Especímenes tipo; Cactaceae; Herbario BA; Argentina.



The relief of natural history collections (NHCs) has been largely recognized for its research and social importance (Lister et al., 2011; Mares, 2009; Suarez & Tsutsui, 2004, Patterson, 2002). The Herbarium of the Argentinian Natural Sciences Museum "Bernardino Rivadavia", BA Herbarium (Holmgren et al., 1990), was initially established in 1853 as the Botanical National Collection of the former Museo Nacional de Historia Natural, representing the first official herbarium of Argentina. Nowadays it preserves more than 150.000 herbarium specimens, mostly representing the main flora of the Argentinian phytogeographical provinces and covering all the vegetated areas of the country, but also collected abroad and obtained by exchange with many national and international botanical institutions. Of all specimen, about 101.500 belong to Vascular Plants, 50.000 to Cellular Plants and 800 represent nomenclatural published type specimens.
Lucien Leon Hauman-Merck (1880-1965) and Carlos Luis Spegazzini (1858-1926) were two of its first botanists researchers, devoted to Argentinian Flora, describing and naming a great number of genera and species. Hauman is author of one of the first and most extensive local catalogues (Hauman, 1917), and collected a number of antique specimen deposited at BA not only from Argentina but also from France. Spegazzini was undoubtedly an enthusiastic plant devoted, considering the large number of publications, extensive works on regional floras and several opuscules, many of them dedicated exclusively to cacti species distributed in Argentina. He had such a predilection for flowering plants in general and the Cactaceae family particularly that he established a great personal herbarium of vascular plants (LPS; Holmgren et al., 1990) constituted by an unknown number of specimens and ca. 700 type specimens, transferred to Museo de La Plata Herbarium (LP) in 1966 (Katinas et al., 2004). Many Spegazzini's specimens are currently found in BA, BAF, BAB, and CORDin Argentina and in Europe and the United States herbarium. His type collection is rather incomplete since most of the Spegazzini's type sheets are missing (Torres at al. 2006; Katinas et al., 2000).
Castellanos and Lelong (1940) refer that the types of the majority of Spegazzini's cacti species were not preserved because they were represented by living specimens, cultivated at home, without labels on the plant itself and he generally made his notes on the spot from the plant. The living specimens cultivated in Spegazzini's residence represent approximately 125 different species (Spegazzini, 1900). Shortly before his death in 1926, Spegazzini donated his collection of live cacti to the Zoological of La Plata. Later, the director of the Zoo at that time, Carlos Marelli, donated some "types" and other of these living specimens to the Argentinian Natural Sciences Museum of Buenos Aires (herbarium BA) (Katinas et al., 2004; Castellanos y Lelong, 1940). It is also well documented in some photographs and on the collection entrance books.
Alberto C. Castellanos was born in Córdoba province, Argentina, on December 11th, 1896. Son of the military Tte. Coronel Julián Castellanos and Isolina Cámara, he was soon encouraged by his mother to study at the Cordoba National University, where he met a still young botanist Federico Kurtz. The latter would have been responsible for initiating Castellanos into the study of plants and introducing him into the experience of collection campaigns (Descole, 1969). Kurtz also encouraged Castellanos to continue his studies on Natural Sciences in Buenos Aires, by contacting him with Dr. C. Hicken, at that time professor at the Exacts and Natural Sciences Faculty from Buenos Aires University.
Castellanos finally obtained his postgraduate degree in 1923 after prolific researches in Cactaceae and Bromeliaceae botanical families. His doctoral thesis appeared two years later, constituting his first publication on Cactaceae (Castellanos, 1925). He worked with almost every vascular plant family, but left and invaluable legacy on regard to an understanding and knowledge of the cacti and bromeliace, being considered a worldwide authority. In 1925, he started working at this Museum as an assistant of Ing. Lucien L. Hauman-Merck, at that time in charge of the Herbarium. The museum was annexed to Natural Sciences Faculty from Buenos Aires University by then. This fact allowed him to serve as both botanist researcher and enthusiastic university teacher. As a teacher he contributed to the education of botanists in several fields beyond phanerogames study: first cryptogamists, geobotanists and paleobotanists in our country were his disciples (Guarrera, 1972).
His first collection campaigns took place all over Argentina in company of Hauman (Singer, 1969), and were aimed to document the national flora. Subsequently, between 1947 and 1976, in collaboration with Roberto Capurro and/or Ofelia Castagnino, it was published in several volumes the first Catálogo de Plantas Vasculares de la Argentina. Thus, his collection specimens started to constitute the main core of the Herbarium. Castellanos prompted the adoption of a systematically organized herbarium (Castellanos, 1936) following the classification system proposed by Engler regarding to botanic Orders and Families and unchanged in following editions (Engler, 1964), and arranging genera within families following De Dalla Torre criteria (Dalla Torre & Harms, 1907), criterium also adopted by most herbaria in the world. While in charge of the Herbarium, as Head of the Botany Department from Argentinian Natural Sciences Museum, the Herbarium obtained the BA acronym according to the International Association of Plant Taxonomists (IAPT), association which he was co-founder. He occupied the Head of Botany Department position until 1948, when became honorary researcher. Then he collaborate and directed the branch of Instituto Miguel Lillo in Buenos Aires city. There, he organized the botanical collections, descriptions and illustrations published in the Genera et Species Plantarum Argentinarum (1945-1955), leaded by Horacio Descole, first global flora from our country.
In 1957, he established in Rio de Janeiro where organized the Museo do Rio de Janeiro Herbarium, referred as Herbário Alberto Castellanos (GUA) which belongs to Instituto Estadual Do Ambiente (INEA), institution responsible for the edition of the scientific Journal "Albertoa". He also teached and contributed with the Jardim Botânico do Rio de Janeiro, city where he finally deceased on the September 5th, 1968.
All over his vast and prolific career, he accounts for more than 26.000 plant specimens collected and more than 100 publications authored, a few of them in collaboration with his wife, Herminia Lelong. He worked at Universidad de Buenos Aires, Universidad Nacional de Tucumán, Universidad de Montevideo and Museo Nacional do Rio de Janeiro. In Brazil, he was awarded with the Orden Nacional do Cruzeiro do Sul and Medalla al Mérito D. Joao VI.
Among his legacy at BA herbarium, there is a vast cactus collection, initiated and developed on his own, almost unique in the remarkable feature of being preserved partially in special conservation liquid. The collection counts with 1.571 specimens, more than a half collected by Castellanos (857 specimens: 54, 6%), some other donated by colleagues or obtained by exchange. Among them all, 60 represent type specimens (3, 8%), most of them from Argentina (52: 88,1%), where Castellanos collected 20 type specimens (33,3 %). There is one specimen from Brasil and another one from Uruguay. From Argentina, Córdoba is the province with more type specimens deposited (11 specimens: 18,3%) followed by La Rioja (8 specimens: 13,3%), San Luis and Catamarca (6 specimens each: 10%).
Castellano's way of work follows this protocole: after field collection, vegetative parts of cactaceas were brought to the Museum (by then located at the old Alsina street building), and cultivated in the yard or at greenhouse. Part of the material corresponding either to vegetative or reproductive stages was then fixed, allowing the collection to account for specimen vegetative parts, flower, fruit and seeds, while preserving the original size and shape (Castellanos, 1940). Fixation utilized formalin (formaldehyde 40%) then diluted once more at 40% in distilled water. The rest of the material was traditionally compressed and dried.
The present article is part of a series of taxonomic articles that included 30 BA type specimens belonging to Potamogetonaceae, Alismataceae, Cyperaceae, Xyridaceae, Erocaulaceae (López, 1988) and 112 to Poaceae family (Arriaga et al., 2001). Here we present a list of the 60 Cactaceae type specimens arranged by the taxon name with author, followed by the publication data (bibliographic citation), classification of type (holotypus, isotypus, paratypus, topotypus, neotypus or lectotypus, typus formae, isotypus varietatis, typus), and all the specimen information included in the original handwritten label. In cases of synonymy, currently accepted names are updated, accompanied by the proper reference to the author and date of the publication that supports the shift. Under observations, ecological information or additional comments written by the author of the description are given. Under notes, some consideration of our own, most regarding the current synonymy, are given.
In agreement with current development of museum-based informatics and its applications in biodiversity analysis (Graham et al., 2004; Edwards et al., 2000), the entire Cactaceae collection was digitized, as part of the broader Digitalization of Biological Collections Project carried out by the MACN. The digitizing process of the BA included a data entry of about 24.000 specimens, and the complete information of each cactus specimen has been achieved in 2008-2009. All data were incorporated to a Microsoft Acces database using Aurora, a special database manager software developed by the MACN(Rodríguez, 2007-2014), and is available online through the Sistema Nacional de Datos Biológicos (SNDB) website (, and the Global Biodiversity Information Facility (GBIF) (
Original names: Original names of taxa appear in italics, followed by author´s standard form and literature data containing the original description. The list of names of species and infraspecific taxa belonging to Cactaceae follows "The International Plant Names Index" (IPNI;, The Flora del Conosur database (IRIS, and Missouri Botanical Garden database (Tropicos, and all other appropriate literature, e.g. publications authored by A. Castellanos where new taxa are described.
Exsiccata: Label transcription of each type specimen is given, including (in order, semicolon spaced): nomenclatural type classification (holotypus, isotypus, paratypus, topotypus, neotypus, etc.), BA number, collector's name and number (if present), collection's date (day/month/year), collection's place information: «country, province, department, area or locality», type of preservation (dried or liquid conserved specimen). The country, political province and department of collection, was added by us when the label lacked that data. When there is not a reference about the location or date of collection, it is referred as missing information.
Observations: Includes collector's or determinavit handwritten comments on the taxon or collection details.
Notes: Relevant consideration of our own and consulted cacti specialist (e.g. R.Kiesling).
When more than one type specimen referring to the same scientific name exist, each of them is listed below the original name (not repeated) and publication (cited once if all of them were described in the same publication).
Current names: Current names of taxa appear in bolded italics, following mainly Zuloaga & Morrone (1999), R. Kiesling publications, Flora del Conosur ( and Tropicos ( databases; with references cited at the end. When the current accepted name is still the original one, this item is omitted and reference correspond to the original name publication.

Notes, abbreviation and acronym list
«»: transcription of the interpreted locality information where specimen was collected.
BA N°: specimen identification number in Museo Argentino de Ciencias Naturales "Bernardino Rivadavia" Herbarium (BA).
BAB: Herbarium from Instituto de Recursos Biológicos, Instituto Nacional de Tecnología Agrícola (INTA).
BAF: Herbarium form Museo de Botánica Juan A. Domínguez, Facultad de Farmacia y Bioquímica, Universidad Nacional de Buenos Aires.
CORD: Herbarium from Museo Botánico Facultad de Ciencias Exactas, Físicas y Naturales, Universidad de Córdoba.
LP: Museo de La Plata Herbarium, Universidad Nacional de La Plata y Museo.
SI: Instituto de Botánica Darwinion Herbarium.
MO: Missouri Botanical Garden Herbarium.
leg.: legit (Latin), collector.
(L): specimen preserved in conservation liquid.
(S): dried-preserved specimen.
(L&S): specimen conserved both in liquid and dried.
Ref.: publication author and date where the current name of the taxon is accepted.
Obs.: Observations.
n. sp., n. spec.: nova species (Latin), new species.
Inter [...] et [...]: Between [...] and [...] (e.g. "inter Portezuelo et La Merced").
m s m: metros sobre el mar (Spanish), meters above sea level.


Cereus silvestrii Speg., Anales del Museo Nacional de Buenos Aires 4: 483. 1905.
Typus- BA Nº 6297; leg.: Spegazzini, Carlos; collection date missing.
«Argentina; Tucumán» (L).
Obs.: Specimen cultivated by C. Spegazzini, traslated before his death to La Plata Zoo and donated to BA, registered in its entrance book 1932 as "Typus Spegazzini" .
= Lobivia silvestrii (Speg.) G.D. Rowley
Ref.: Rausch, W. 1986.

Echinopsis oxygona (Link.) Zucc. ex Pfeiffer & Otto, Abbild. Beschr. Cact. 1: t. 4.1838.
typus- BA Nº 6298; collector and collection date missing. (L).
Obs.: Specimen cultivated by C. Spegazzini, traslated before his death to La Plata Zoo and donated to BA, registered in its entrance book 1932 as "Typus Spegazzini" .
Note: Species not included in IPNI.

Gymnocalycium andreae (Boed.) Backeb. subsp. matznetteri Rausch, Gymnocalycium 13(4): 379. 2000.
Holotypus- BA Nº 80542; leg.: Rausch, W. 567a, collection date missing.
«Argentina; Córdoba; Sierra Grande, Los 5
Gigantes» (S).

Gymnocalycium andreae (Boed.) Backeb. var. longispinum Rausch, Gymnocalycium 13(4): 378. 2000.
Holotypus- BA Nº 80547; leg.: Rausch, W. WR108, collection date missing.
«Argentina; Córdoba; Sierra Grande, El Cóndor» (L).
Obs.: Existe en herbario una diapositiva.

Gymnocalycium andreae (Boed.) Backeb. var. fechseri H. Till, Gymnocalycium 13(4): 378. 2000.
Typusvar. - BA Nº 80546; leg.: Fechser, H. (H.Till 1308); 1974; collection date missing. (L).

Gymnocalycium bicolor Schütz
LECTOTYPUS- BA Nº 28/252; leg.: Castellanos, A.; 14/02/1928.
«Argentina; Córdoba; Cruz del Eje» (L).

Lectotypification designed by Řepka R. (2014): Typification of the name Gymnocalycium bicolor Schütz with taxonomic-nomenclatorical notes. Bradleya 32: xx-xx (in press).
Gymnocalycium borthii Koop. ex H. Till var. viridis Neuhuber, Gymnocalycium 20(1): 705. 2007.
Holotypus- BA Nº 92224; leg.: Neuhuber, G. GN89-85; 18/12/1989.
«Argentina; San Luis; Cerros del Rosario, 1000 m s.m» (S)

Holotypus- BA Nº 92217; leg.: Neuhuber, G. GN89-85/476; 18/12/1989.
«Argentina; San Luis; Cerros del Rosario; 1000 msm» (L)
Note: According to IRIS (SI Herbarium) and TROPICOS (MOHerbarium) taxonomic databases, Gymnocalycium borthii Koop. ex H. Till is synonym of the accepted name Gymnocalycium gibbosum (Haw.) Pfeiff. ex Mittler, but not any viridis Neuhuber variety is registered
Gymnocalycium capillense (Schick) Hosseus., Revist. Cent. Estud. Farm. Córdoba reimpr 2(6): 16. 1926.
Neotypus- BA Nº 92216; leg.: Fechser, H. s.n.; 1961. Till,H. 2003. Gymnocalycium 16 (2): 517.
«Argentina» (S)
Ref.: Metzing, D. et al. 1995; Meregalli, M. 1985.

Gymnocalycium capillense (Schick) Hosseus. var. sigelianum H.Till, Gymnocalycium 16(1): 497. 2003.
NEOTYPUS - BA Nº 92214; leg.: Till, H. HT58; collection place and date missing. (L)
Gymnocalycium castellanosii Backeb. var. rigidum H. Till, Gymnocalycium 19(2): 670. 2006.
HolotYpUS - BA Nº 92218; leg.: Till, H. 900;16/09/1961.
«Argentina; La Rioja; Belgrano; NW del Departamento Belgrano» (S)
Gymnocalycium coloradense F. Berger, Gymnocalycium 19(4): 693. 2006.
HolotYpUS - BA Nº 92221; leg.: Berger, F. 97-72/291; 09/11/1997.
«Argentina; La Rioja; Independencia; Sierra de los Colorados, in parte australi, 700-900 m s m.» (S)
Gymnocalycium 19(4): 691. 2006.
Holotypus- BA Nº 92222; leg.: Berger, F. Be97-72/291; 09/11/1997.
«Argentina; La Rioja; Independencia; Sierra de los Colorados, in parte australi; 700-900 m s m» (L)
Gymnocalycium 19(4): 691. 2006.
PARAtypus- BA Nº 92272; leg.: Berger, F. Be01-271/1237; 17/11/2001.
«Argentina; La Rioja; Independencia; Sierra de los Colorados, in parte australi; 700-900 m s m» (L)
Gymnocalycium delaetii Hosseus f. antherosacos Neuhuber
NEOtYpUS for. - BA Nº 80545 ; leg.: Neuhuber, G. GN88-41/72 (WU1312); Collection date missing.
«Argentina; Salta; Alemania» (L)
Note: According to IPNI, the type details are:

Gymnocalycium delaetii (K.Schum.) Hosseus fo. antherosacos F.Ritter ex Neuhuber, Gymnocalycium 14 (4): 423. 2001.
G.Neuhuber GN88-41; 02/05/1988; «Argentina; Salta; Australo-occidentalier urbis Alemania in solo arenoso Sub fructibus, 1400 m».
Gymnocalycium erolesii Neuhuber & Bertch Gymnocalycium 15: 475. 2002.
typus- BA Nº 81420; Collector name missing; 24/11/2000.
«Argentina; Santa Fe; NE Vera» (L)
Obs.#1: typus pars 2. Pfl-Nummer: PE0025. Fundort: NE Vera, Santa Fe.
Obs.#2: typus pars 2. Pfl-Nummer: PE0025. Fundort: between Vera and Malabrigo, Santa Fe.
Note: It's only one sample in two bottles, both samples are the same PE0025

= Gymnocalycium schroederianum Osten subsp. boessii R. Kiesling, Marchesi & O. Ferrari
Ref.: Kiesling, R., Marchesi, E. & Ferrari, O. 2002.
Note: According to IPNI, the BA holotype details are:

Gymnocalycium erolesii Neuhuber & C.A.L.Bercht, Gymnocalycium 15(4): 475. 2002.
Holotypus; leg.: Eroles, P. PE00-25; 24/11/2000.
«Argentina; Santa Fe; inter urbium Vera et Malabrigo, km 742 viae "ruta internacional no. 11"»
Gymnocalycium gaponii Neuhuber, Gymnocalycium 14(4): 410. 2001.
tYpUS - BA Nº 80543; leg.: Neuhuber, G. 91-362/1222.
«Argentina; Córdoba; San Lorenzo» (L)
Note: According to IPNI, the holotypus details are:

Gymnocalycium gaponii Neuhuber, Gymnocalycium 14(3): 410. 2001.
leg.: G.J.A. Neuhuber 91-362/1222; 07/12/1991; «Argentina; Córdoba; prope Villa Rafael Benegas, 900m»
ParatYPUS - BA Nº 80544; leg.: Neuhuber, G. 96-850/2783; 25/10/1946.
«Argentina; Córdoba; San Lorenzo, Sierras de San Luis» (L)
Note: Collection date probably wrong.

Gymnocalycium immemoratum A. Cast. & Lelong, Lilloa 4(2): 195-196.1939.
Holotypus- BA Nº 30526; leg.: Castellanos, A. & Lelong; 06/01/1939.
«Argentina; Córdoba; Punilla; Capilla del Monte, El Zapato» (L y S)
Obs.: Determinavit R. Kiesling et OF/88: G. valnicekianum.

= Gymnocalycium mostii (Gürke) Britton & Rose
Ref.: Till, H. & Amerhauser, H. 2002; Metzing, D. et al. 1995; Meregalli, M. 1985; Britton, N.L. & Rose, J.N. 1918.

Gymnocalycium jochumii var. jugum Neuhuber, Gymnocalycium 18(1): 606, figs. 14-15. 2005.
HolotYpUS - BA Nº 92219; leg.: Neuhuber, G. GN96-891/2931; 09/10/1996.
«Argentina; La Rioja; Chilecito; Sierra de Sañogasta; 2300 msm» (L)
Obs.: L.Nr.: 3299. Fundort: Southern end of Sa. de Sañogasta, L.R.

Gymnocalycium multiflorum (Hook) Britton & Rose var. ourselianum Cels ex H.Till & W.Till. Gymnocalycium 20(2): 719, figs. 13-16. 2007.
Holotypus- BA Nº 92220; leg.: Rausch, W. s.n. ex coll. H.Till 3541; 10/06/1994.
«Argentina; Loco exacto ignoto» (S)
Note: According to IPNI, the taxa is Gymnocalycium multiflorum Britton & Rose var. ourselianum H.Till & W.Till,
Ref.: Till, H. & Till, W., 2007.
Obs.: this var. is not considered as valid in Tropicos.

Gymnocalycium nigriareolatum Backeb. var. densispinum Backeb. ex H.Till, Till, H. Gymnocalycium 11(3): 255. 1998.
Holotypusvar. - BA Nº 79588; leg.: Fechser, H. s.n.; 1963.
«Argentina; Sine loco exacto» (S)
Obs.: 1998 cult. in coll H.Till #HT607.

= Gymnocalycium hybopleurum (K. Schum.) Backeb. Ref.: Till, H. & Till, W. 1995, 1994; Backeberg, C. & Kunth, F.M. 1935.
Note: According to IPNI, type details are:
leg.: H.Till HT607; sine loco exacto, coll. H. Fechser s.n., 1963, cult. in coll. H. Till.
Remarks: Backeberg (Cactaceae 3: 1759. 1959) published this name invalidly. The type collection, originally collected by H. Fechser in 1963, flowered in cultivation.

Gymnocalycium nigriareolatum Backeb. var. simoi H. Till, Gymnocalycium 11(3): 258. 1998.
Holotypusvar. - BA Nº 79589; leg.: Borth, H. BO126; 1972.
«Argentina; Catamarca; Paclín; Palo Labrado, inter Portezuelo et La Merced» (S)
Obs.: 1998 cult. in coll. H.Till #HT1227

Gymnocalycium nigriareolatum Backeb. f. carmineum H. Till, Gymnocalycium 11(3): 255. 1998.
Holotypusfor. - BA Nº 79587; leg.: Amerhauser, H. HA255-2; 08/10/1989
«Argentina; Catamarca; Western slopes of the Sierra de Ancasti 600-700 m s m.» (S)

Gymnocalycium ragonesei A. Cast., Lilloa 23: 5.1950.
Isotypus- BA Nº 80073; leg.: Ragonese, A.; 7 Fadel et al.: Cactaceae type specimens at BA
«Argentina; Catamarca; Salinas Grandes, entre Km 969 y Totoralejos» (L)
Obs.: Specimen LIL16120 duplo

[Gymnocalycium rhodantherum (Boed.) H. Till], Gymnocalycium 18(1): 602. 2005.
Neotypus- BA Nº 92215; leg.: Till, H. 87-67; 18/10/1987.
«Argentina; La Rioja; Famatina; km 525 entlang der Ruta Nacional südlich Pituil, 1250 m s m.» (S)
Obs.: Inval. Art. 9.20. Gymnocalycium 20(1): IV (2007)!.
Note: R. Kiesling (2005) considers this a doubtful taxon.

[Gymnocalycium rhodantherum (Boed.) H. Till var. cinerascens H. Till], Gymnocalycium 17(3): 584. 2004.
Holotypus- BA Nº 92213; leg.: Till, H. 90-308; 07/02/1990.
«Argentina; La Rioja; Prope Carrizal; 1800 m.s.m.» (S)
Note: Kiesling (2005) considers this a doubtful taxa.

Gymnocalycium schickendantzii subsp. bergeri Neuhuber, Gymnocalycium 14(4): 420. 2001.
HolotYpUS - BA Nº 80541; leg.: Neuhuber, G. GN90-308; 25/12/1990. (L y S)
Obs.: Specimen (L): «Argentina; Salta; In silvis collium circa Balboa, 700 m s m.» (L)
Dried specimen (S): 1391 GN90-308 «Los Baños, Salta».
Note: part of the specimen was dried and the other part conserved in liquid.
Ref.: Neuhuber, G.J.A, 2001b.

Gymnocalycium schroederianum Osten, Anales Museo Nacional Montevideo, ser. 2 5: 60, pl. 49–50. 1941.
Isotypus- BA Nº 24/1133; leg.: Schröder, J. s.n.; 01/04/1922.
«Uruguay; Río Negro; Cerca de Nueva Mehlem, en limo pampeano a orillas del Río Uruguay» (S)
Obs.: in limo pampeano ad ripam fluminis Uruguay, prope Nueva Mehlem.
Ref.: Kiesling, R. 2005; Kiesling, R. 1987; Osten, C. 1941.

Gymnocalycium taningaense Piltz subsp. fuschilloi Neuhuber, Gymnocalycium 20(1): 705. 2007.
Holotypus- BA Nº 92223; leg.: Neuhuber, G. GN90-240; 15/12/1990.
«Argentina; Córdoba; Meridionaliter de Arroyo San Antonio; 630 msm» (S)
Hariota epiphylloides var. bradei Porto & A. Cast., Rodriguésia (5)14: 354. 1941.
IsotYpUS var. - BA Nº 20308; leg.: Brade 20; 20/07/1937.
«Brasil; São Paulo; Serra Bocaina, sertão do Rio Vermelho» (L)
Obs.: R.n. Jardim Bot. 8451

= Hatiora epiphylloides subsp. bradei (Porto & A. Cast.) Barthlott & N.P.Taylor
Ref.: Barthlott & Taylor, 1995

Lobivia schreiteri A. Cast., Monatsschr. Deutsch. Kakteen.-Ges. 2(3-4):59.1930.
Holotypus- BA Nº 29/41; leg.: Castellanos, A.; 20/07/1929.
«Argentina; Tucumán; Portezuela de la Ciénaga» (L y S)
Maihuenia valentinii Speg., Anales Mus. Nac.Buenos Aires 7 (ser. 2, tomo 4): 289. 1902.
Isotypus- BA Nº 17397; leg.: Valentini, Juan 48; 11/11/1897.
«Argentina; Chubut; Rawson; Trelew, Cima Alta» (S)
Obs.: «Terr. del Chubut. Trelew. Cima Alta».

= Maihuenia patagonica (Phil.) Britton & Rose
Ref.: Leuenberger, B. E. 1997; Kiesling, R. 1988; Britton, N.L. & Rose, J.N. 1919.
Notocactus mammulosus (Lem.) A. Berger var. pampeanus (Speg.) A. Cast. & Lelong, Descole Gen. Sp. Pl. Argent. 1: 100. 1943.
Topotypus- BA Nº 30525; leg.: Castellanos, A. y H. Lelong; 06/01/1939. «Argentina; Córdoba; Capilla del Monte, El Zapato. (L)

= Parodia submammulosa (Lem.) R. Kiesling subsp. submammulosa
Ref.: Kiesling, R. 1995.

Opuntia anacantha Speg., Bull. Mus. Hist. Nat. (Paris) 10: 391. 1904; Anal. Mus. Nac. Buenos Aires, 11: 513. 1905.
ISOTypus- BA Nº 6295; Collector name, and collection date missing.
«Argentina, Chaco Austral» (L)
Note: Specimen collected in Chaco Austral region and then cultivated by C. Spegazzini, traslated before his death to La Plata Zoo and donated
to BA, registered in its entrance book 1932 as "Typus Spegazzini" Ref.: Kiesling, R. 1998.

Opuntia geometrica A. Cast., Kakteenkunde 9: 192, 1934.
HOLOTyPus- BA Nº 30/748; leg.: Castellanos, A.; 24/01/1930.
«Argentina; Catamarca; Tinogasta; Angostura de Guanchín» (L & S)

= Tephrocactus geometricus (A. Cast.) Backeb.
Ref.: Kiesling, R. 1984
Opuntia montevidensis Speg., Anales Mus. Nac. Buenos Aires ser. 3, 4: 515.1905.
Typus- BA Nº 6299; Collector name, collection date and place missing. (L)
Note: Specimen collected and then cultivated by C. Spegazzini, traslated before his death to La Plata Zoo and donated to BA, registered in its entrance book 1932 as "Typus Spegazzini".

= Opuntia aurantiaca Lindl.
Ref.: Kiesling, R. 2005.
Opuntia paediophila A. Cast., Lilloa 23: 7.1950.
Holotypus- BA Nº 30/759; leg.: Castellanos, A.; 17/02/1930.
«Argentina; La Rioja; San Blas de los Sauces; Alpasinche» (L)
Paratypus- BA Nº 80074; leg.: Ragonese, A.; 19/10/1946.
«Argentina; Córdoba; Cruz del Eje; Salinas Grandes, Mansilla, Km 907» (L)
Obs.: Specimen LIL16122 DUPLO
ParatYpUS - BA Nº 28/643; leg.: Gómez; 07/1928.
«Argentina; La Rioja; Rosario Vera Peñaloza; Chepes» (L)

= Tephrocactus aoracanthus (Lem.) Lem.
Ref.: Kiesling, R. 1984.
Opuntia puelchana A. Cast., Physis 9: 101. 1928.
HOLOTYpUS - BA Nº 27/195; leg.: Castellanos, A.; 10/01/1927.
«Argentina; La Pampa; Lihuel Calel; Pampa Central: Lihué-Calel» (L)

= Cylindropuntia tunicata (Lehm.) F.M. Knuth
Ref.: Hoffmann, A. E. & Walter, H. E. 2004; Backeberg, C. 1958.
Opuntia ruiz-lealii A. Cast., Lilloa 9: 213. 1943.
TopotYpUS - BA Nº 36992; leg.: Castellanos, A.; 06/01/1941.
«Argentina; Mendoza; Las Heras; Valle de Uspallata» (L)

= Puna clavarioides (Pfeiff.) R. Kiesling
Ref.: Kiesling, R. 1982.
Opuntia salagria A. Cast., Lilloa 27: 85-89, 1953.
SINTYpUS - BA Nº 27/196; leg.: Castellanos, A.; 05/01/1927.
«Argentina; La Pampa; Capital; Pampa Central, Santa Rosa, El Médano» (L)
SINTYPUS - BA Nº 26/2363; leg.: Castellanos, A.; 05/12/1926.
«Argentina; Buenos Aires; Campana; Campana, en las Barrancas» (L)
SINTYPUS - BA Nº 26/1849; leg.: Castellanos, A.; 04-07/10/1926.
«Argentina; Santa Fe; Rosario; Rosario, en los barrancos» (L)
SINTYPUS - BA Nº 25/855; leg.: Castellanos, A.; 25/02/1925.
«Argentina; San Luis; Belgrano; Nogolí, Paso Ancho» (L)
SINTYPUS - BA Nº 25/2619; leg.: Castellanos, A. 843; 12/11/1925.
«Argentina; San Luis; Chacabuco; Renca» (L)
SINTYPUS - BA Nº 25/2615; leg.: Castellanos, A.; 13/11/1925.
«Argentina; San Luis; Chacabuco; Renca» (L)
SINTYPUS - BA Nº 32594; leg.: Fortuna; 11/1939.
«Argentina; La Pampa; Loventué; Pampa Central, Loventuel» (L)
SINTYPUS - BA Nº 33033; leg.: Ragonese, A.; 20/03/1940.
«Argentina; Santa Fe; 9 de Julio; Villa Minetti» (L)
SINTYPUS - BA Nº 54768; leg.: Castellanos, A.; 20/11/1951.
«Argentina; Buenos Aires; Campana; Campana» (L-S)
Obs.: F. purpúreos.
SINTYPUS - BA Nº 30541; leg.: Castellanos, A.; 29/01/1939.
«Argentina; Córdoba; San Javier; Pedanía La Paz, Quebrada del Cristal de Roca» (L)
SINTYPUS - BA Nº 24/883; leg.: Castellanos, A.; 02/1924.
«Argentina; Córdoba; Valle de los Reartes» (L)
SINTYPUS - BA Nº 24/1344; leg.: Castellanos, A.; 21/07/1924.
«Argentina; Córdoba; Punilla; Dique San Roque» (L)
TOPOTYPUS - BA Nº 29708; leg.: Castellanos, A. 19534; 20/11/1951.
«Argentina; Buenos Aires; Campana; Barrancas de Campana» (S)
= Opuntia megapotamica Arechav.
Ref.: Kiesling, R. 2005.
Opuntia weberi Speg. var. dispar A. Cast. & Lelong in Jahrb. Deutsch. Kakteen-Ges. l: 51. 1935/36.
MATERIAL HISTÓRICO(utilizado para la ilustración en la ampliación de la descripción de la variedad, realizada por Castellanos & Lelong (1944)Lilloa X: 470
- BA Nº 13514; leg.: Peirano; 27/12/1933.
«Argentina; Catamarca; Santa María; Andalhualá» (S)
Obs.: Fruto rojo.

= Tephrocactus weberi (Speg.) Backeb. var. weberi
Ref.: Kiesling, R. 1984; Backeberg, C. & Knuth, F.M., 1935.
Pterocactus araucanus A. Cast., Rev. Fac. C. Agr. Univ. Nac. Cuyo. 8(2): 3-5 1960-61.
HolotYPUS -BA Nº 80061; leg.: Castellanos, A. 16812; 14/02/1945.
«Argentina; Chubut; Mesetas cerca del Río Hualjaina» (L).
Obs.: Specimen LIL15301 DUPLO
Rev. Fac. C. Agr. Univ. Nac. de Cuyo. 8(2): 3 - 13. 1960-61.
ParatYpUS - BA Nº 13471; leg.: Ragonese, A.; 01/1935.
«Argentina; Neuquén; Zapala; Zapala» (L).
Ref.: Kiesling, R. 1982; Castellanos, A. 1960.


Many thanks are given to Roberto Kiesling for critical reading of the manuscript and synonymy revision, personal notes on names, distribution and reference information. We also acknowledge Lic. Cinthia Susic Martín for partial data entry. The digitizing project was supported by William Foundation, the Japan International Cooperation Agency (JICA) and CONICET.


1. Arriaga, M.O., R.A. Gómez Cadret & Ma. D. Monteros. 2001. Catálogo de tipos depositados en el herbario del Museo Argentino de Ciencias Naturales "Bernardino Rivadavia" (BA).II. Monocotiledóneas: Poaceae. Rev. Mus. Argentino Cienc. Nat. 3(2):135-146.         [ Links ]

2. Backeberg. C. 1934. Blätter für Kakteenforschung. (5): 8.         [ Links ]

3. Backeberg. C. 1958. Die Cactaceae I. Fischer Verlag. Jena.         [ Links ]

4. Backeberg, C. & F.M. Knuth. 1935 [1936]. Kaktus-ABC: 1-432.         [ Links ]

5. Barthlott, W. & N. P. Taylor. 1995. Notes towards a monograph of Rhipsalideae (Cactaceae). Bradleya 13: 43 – 79.

6. Berger, F. 2006. Ergebnisse langjahriger Pflanzenstudien und Beobachtungen an den heimatlichen Standorten. Eine beachtenswerte Sippe aus der Sierra de Los Colorados, Prov. La Rioja, Argentinien. Teil 4: Gymnocalycium coloradense F.Berger spec. nov. Gymnocalycium 19(4): 691-696.         [ Links ]

7. Britton, N.L & N.J. Rose. 1918. Gymnocalycium mostii (Gürke) Britton & Rose. Addisonia 3: 5, pl. 83.         [ Links ]

8. Britton, N.L. & J.N. Rose. 1919. The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 41.         [ Links ]

9. Castellanos, A.C. 1925. Rhipsalis argentinas (Cactaceas). Anales Mus. Nac. Buenos Aires 32: 477-500.         [ Links ]

10. Castellanos, A.C. 1928. Opuntia puelcheana Cast. n. sp. Physis. Revista de la Sociedad Argentina de Ciencias Naturales 9: 101.         [ Links ]

11. Castellanos, A.C. 1930. Lobivia schreiteri Castellanos n.sp. Monatsschr. Deutsch. Kakteen.-Ges. 2 (3-4): 59-60.         [ Links ]

12. Castellanos, A.C. 1934. Opuntia geometrica Castellanos, n.sp. Kakteenkunde, Berlin. 9:172. Illustr.         [ Links ]

13. Castellanos, A.C. 1943. Opuntia ruiz-lealii Castell. nova sp. Lilloa 9: 211-213.         [ Links ]

14. Castellanos, A.C. 1950. Dos especies nuevas de Cactáceas Argentinas. Lilloa. 23: 5-13.         [ Links ]

15. Castellanos, A.C. 1953. Noticias sobre Opuntia (Cactaceae). Lilloa 27: 81-89.         [ Links ]

16. Castellanos, A.C. & H.V. Lelong. 1939. Notas fanerogámicas. Gymnocalycium immemoratum Castellanos n. sp. Lilloa 4(2): 195-196.         [ Links ]

17. Castellanos, A.C. & H.V. Lelong. 1940. The Cactus family in the Museum of Natural Sciences of Buenos Aires. Cactus & Succulent Journal, Pasadena. 12(3): 49-51.         [ Links ]

18. Campos Porto, P. & A.C. Castellanos. 1941. Hariotae novae brasiliensis. Rodriguèsia, R. de Janeiro, 5 (14): 353-354, pl.1-5.         [ Links ]

19. Castellanos, A.C& H.V. Lelong. 1943. Opuntiales vel Cactales. H. Descole (ed.). Gen. Sp. Pl. Argent. 1: 47-142.         [ Links ]

20. Castellanos, A.C& H. V. Lelong. 1944. Opuntia weberi Spegazzini var. dispar Castell. et Lelong. Lilloa 10: 469-470.         [ Links ]

21. Castellanos, A.C& H.V. Lelong. 1944. Una nueva especie de Opuntia. Lilloa, 10:395-402.         [ Links ]

22. De Dalla Torre, C.G. & H. Harms. 1907. Genera Siphonogamarum ad Systema Englerianum Conscripta. Lipsiae, 921 pgs.         [ Links ]

23. Descole, H.R. 1969. Alberto Castellanos 1896-1968. Revista de la Facultad de Ciencias Agrarias de Mendoza 15(1): 168-178.         [ Links ]

24. Eggli, U. & B.E. Leuenberger. 1998. On colour forms of Opuntia weberi (Cactaceae) with notes on the typification of the name. Willdenowia 28: 175-180.         [ Links ]

25. Edwards, J.L., M.A. Lane & E.S. Nielsen. 2000. Interoperability of biodiversity databases: biodiversity information on every desktop. Science 289, 2312–2314.

26. Engler, H.G.A. 1964. Syllabus der Pflanzenfamilien, H.Melchior (Ed.), 12th edition, Vol.2. Borntraeger, Berlin.         [ Links ]

27. Guarrera, S. en Kiesling, R. (Edit.) 1972. Alberto Castellanos. Recopilación de sus publicaciones sobre Cactáceas. Círculo de coleccionistas de cactus y crasas de la República Argentina.         [ Links ]

28. Hoffmann, A. E. & H.E. Walter. 2004. Una Guía para la identificación de los cactos que crecen en el país. 2da. Edición. Cactáceas en la flora silvestre de Chile. 10-307. Fundación Claudio Gay, Santiago de Chile.         [ Links ]

29. Holmgren, P.K., N.H. Holmgren & L.C. Barnett. 1990. Index Herbariorum. Part I: The herbaria of the world. 8th edition. New York: New York Botanical Garden, 693 pp. [Regnum Veg. 120] (Current electronic version:        [ Links ]

30. Katinas, L., D.G. Gutiérrez & S.S. Torres Robles. 2000. Carlos Spegazzini (1858-1926): Travels and botanical work on vascular plants. Ann. Missouri Bot. Gard. 87: 183-202.         [ Links ]

31. Katinas, L., D.G. Gutiérrez & S.S. Torres Robles. 2004. Type material of Carlos L. Spegazzini in the Museo de La Plata Herbarium (LP), Argentina. III. Cactaceae. Darwiniana 42(1-4): Castellanos, A. & H.V. Lelong 177-200.         [ Links ]

32. Kiesling, R. 1982. Puna, un género nuevo de Opuntioideae (Cactaceae). Hickenia. 1: 289-294.         [ Links ]

33. Kiesling, R. 1987. Two new subspecies of Gymnocalycium schroederianum. Cact. Succ. J. (Los Angeles). 59: 44-49.         [ Links ]

34. Kiesling, R. 1984. Estudios en Cactaceae de Argentina: Maihueniopsis, Tephrocactus y géneros afines (Opuntioideae). Darwiniana. 25: 171-215.         [ Links ]

35. Kiesling, R. 1995. Argentine Notocacti of the genus Parodia. Cactus and Succulent Journal 67 (1): 14-15.         [ Links ]

36. Kiesling, R. 1998. Nota sobre Opuntia anacantha Speg. (Cactaceae) para la flora de Paraguay y Argentina. Candollea 53: 471-476.         [ Links ]

37. Kiesling, R. 2005. Cactaceae en Flora de Entre Ríos, Colecc. Ci. Inst. Nac. Tecnol. Agropecu. 4: 401-444.         [ Links ]

38. Kiesling, R., E. Marchesi & O. Ferrari. 2002. Gymnocalycium schroederianum subsp. boessii, subsp. nov. Kakteen Sukk. 53(9): 225-232.         [ Links ]

39. Lister, A.M., S.J. Brooks, P.B. Fenberg, A.G. Glover, K.E. James, K.G. Johnson, A.M. Lister, E. Michel, B. Okamura, M. Spencer, J.R. Stewart, J.A. Todd, E. Valsami-Jones & J. Young. 2011. Natural history collections as sources of longterm datasets. Trends Ecol. Evol. 26, 153–154.

40. López, M.E. 1988. Catálogo de los tipos depositados en el Herbario del Museo Argentino de Ciencias Naturales "Bernardino Rivadavia" (BA). I. Monocotiledóneas: Potamogetonaceae, Alismataceae, Cyperaceae, Xyridaceae, Eriocaulaceae. Rev. Mus. Argentino Cienc. Nat. "Bernardino Rivadavia", Botánica 7(1): 1-8.         [ Links ]

41. Mares, M.A. 2009. Natural Science Collections: America's irreplaceable resource. BioScience 59:544-545.         [ Links ]

42. Meregalli, M. 1985. Il genere Gymnocalycium Pfeiffer. Piante Grasse. 5: 5-63.         [ Links ]

43. Metzing, D., M. Meregalli & R. Kiesling. 1995. An annotated checklist of the genus Gymnocalycium Pfeiffer ex Mittler (Cactaceae). Allionia. 33: 181-228.         [ Links ]

44. Neuhuber, G.J.A. 2001a. Gymnocalycium gaponii, eine neue Art von der Westseite der Sierra Grande in Argentinien. Gymnocalycium 14. (3): 409-412        [ Links ]

45. Neuhuber, G.J.A. 2001b. Ein Neufund in der Provinz Salta: Gymnocalycium schickendantzii subsp. bergeri und die Validierung und Neubewertung von Ritter's Gymnocalycium antherosacos als Gymnocalycium delaetii fa. antherosacos. Gymnocalycium. 14(4): 417-424        [ Links ]

46. Neuhuber, G.J.A. 2005. Neufunde in der sudlichen Sierra de Sanogasta: Gymnocalycium jochumii und Gymnocalycium jochumii var. jugum. Gymnocalycium. 18(1):603-610.         [ Links ]

47. Neuhuber, G.J.A. 2007. Zur geografischen Radiation des Gymnocalycium borthii und seiner Verwandten. Gymnocalycium 20. (1): 697-708.         [ Links ]

48. Neuhuber, G.J.A & C.A.L. Bercht. 2002. Gymnocalycium erolesii, ein interessanter Fund aus der Provinz Santa Fe. Gymnocalycium 15. (4): 473-478.         [ Links ]

49. Osten, C. 1941. Notas sobre Cactáceas. Anales Museo Nacional Montevideo, ser. 2 (5): 60, pl. 49–50.

50. Patterson, B.D. 2002. On the continuing need for scientific collecting of mammals. Mastozoología Neotropical 9, 253–262.

51. Pin, A.B. 1998. Nota sobre dos especies de Brasiliopuntia (Cactaceae). B. schulzii y B. brasiliensis. Rojasiana. 4(2): 144-163.         [ Links ]

52. Porto, P.C. & A. Castellanos. 1941. Hariotae novae brasiliensis. Rodriguésia (5)14: 353-354.         [ Links ]

53. Rausch, W. 1986. Lobivia 85. Wien.: Rudolf Herzig 160 pp.         [ Links ]

54. Rausch, W. 2000. Drei neue Taxa aus der Gruppe des Gymnocalycium andreae. Gymnocalycium 13 (4): 377–380.

55. Rodríguez, D. 2007-2014. Aurora. Aplicación para el manejo de bases de datos de colecciones biológicas. Museo Argentino de Ciencias Naturales «Bernardino Rivadavia», CONICET.         [ Links ]

56. Singer, R. 1969. Alberto Castellanos 1896-1968, Taxón 18:308-309.         [ Links ]

57. Spegazzini, C. 1900. Notas botánicas. Anales Soc. Ci. Argent. 49: 122-125.         [ Links ]

58. Spegazzini, C. 1902. Nova adenda ad Floram Patagonicam. Anales Mus. Nac. Buenos Aires 7 (ser.2, tomo 4): 135-308.         [ Links ]

59. Spegazzini, C. 1905. Cactacearum platensium tentamen. Anales Mus. Nac. Buenos Aires 11 (ser. 3, tomo 4): 477-521.         [ Links ]

60. Řepka R. (2014): Typification of the name Gymnocalycium bicolor Schütz with taxonomicnomenclatorical notes. – Bradleya 32: xx-xx (in press).

61. Till, H. 1998. Zur Identitat von G. nigriareolatum Backeberg und seinen Formen. Gymnocalycium 11 (3): 251-258.         [ Links ]

62. Till, H. 2004. Bemerkungen zu Gymnocalycium rhodantherum und den mit ihm verwandten Taxa. 2. Zur Identitat von Gymnocalylcium guanchinense Schutz. Gymnocalycium 17 (3): 585-588.         [ Links ]

63. Till, H. 2005. Bemerkungen zu Gymnocalycium rhodantherum und den mit ihm verwandten Taxa. 5.) Gymnocalycium alboareolatum Rausch. Gymnocalycium 18 (1): 611-614.         [ Links ]

64. Till, H. 2006. Ergebnisse langjahriger Pflanzenstudien und Beobachtungen an den heimatlichen Standorten; Teil 1. Gymnocalycium castellanosii s. str. Gymnocalycium 19. (2): 663-670.         [ Links ]

65. Till, H. & H. Amerhauser. 2002. Revision des Gymnocalycium mostii-Aggregates Teil II: Gymnocalycium valnicekianum und seine Formen. Gymnocalycium. 15(2): 445-452.         [ Links ]

66. Till, H & W. Rausch. 2000. Drei neue Taxa aus der Gruppe des Gymnocalycium andreae. Gymnocalycium 13. (4): 377-380.         [ Links ]

67. Till, H. & W. Till. 1994. Gymnocalycium hybopleurum 1. Gymnocalycium 7: 135-140.         [ Links ]

68. Till, H. & W. Till. 1995. Gymnocalycium hybopleurum 2. Gymnocalycium 8: 141-146.         [ Links ]

69. Till, H. & W. Till. 2007. Ist Echinocactus ourselianus Cels ex Salm-Dyck wirklich ein Synonym von Gymnocalycium multiflorum (Hook.) Britton & Rose. Gymnocalycium 20. (2): 715-720.         [ Links ]

70. Torres, M.A., M.O. Arriaga & Ma.D. Montero. 2006. Catálogo de tipos de Poaceae del Herbario del Museo de La Plata (LP). Contr. Mus. Argent. Cs. Nat. "Bernardino Rivadavia" 3: 1-87.         [ Links ]

71. Zuloaga, F.O. & O. Morrone (eds.). 1999. Catálogo de las plantas vasculares de la República Argentina. II. Acanthaceae-Euphorbiaceae (Dicotyledoneae). Mono        [ Links ]

Recibido: 23-IV-2014
Aceptado: 11-XII-2014

Creative Commons License All the contents of this journal, except where otherwise noted, is licensed under a Creative Commons Attribution License